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Curriculum Vitae
Joy Halverson, DVM, MPVM
Director and Senior Scientist
QuestGen Forensics
 
 
 

Professional Experience

12/01-present   Director and Senior Scientist , QuestGen Forensics, LLC, 1046 Olive Drive Suite1, Davis, CA
4/96-11/01 Scientist, Celera AgGen, 1756 Picasso Avenue, Davis, CA
1/89-4/96 President, Zoogen, Inc. 1756 Picasso Ave, Davis, CA
1/87-12/88 Post-doctoral Researcher, University of California, Davis, CA
7/81-12/86 Veterinary clinician, veterinary hospitals in Canberra, Australia and Sacramento, CA

Education

Year   Degree Institution Specialization
1985 Masters University of California, Davis, CA Veterinary Epidemiology
1981 Doctorate   University of California, Davis, CA Veterinary Medicine
1976 Bachelor University of California, San Diego, CA   Biology

Professional Memberships

American Academy of Forensic Science
American Veterinary Medical Association
Association of Avian Veterinarians
International Society of Animal Genetics

Publications (Return to Top)
J. Halverson, C. Basten. 2005. A PCR multiplex and database for forensic DNA identification of dogs. Journal of Forensic Science 50(2):352-363.
S. Denise, E. Johnston, J. Halverson, K. Marshall, D. Rosenfeld, S. Mckenna, T. Sharp, J. Edwards. 2004.Power of exclusion for parentage verification and probability of match for identity in American Kennel Club breeds using seventeen canine microsatellite markers. Animal Genetics. 35(1): 14-17.
C. Vila, C. Walker, A. Sundqvist, O. Flagstad, Z. Andersone, A. Casulli, I. Kojola, H. Valdman, J. Halverson, H. Ellengren. 2003. Combined use of maternal paternal and bi-parental genetic markers for the identification of wolf-dog hybrids. Heredity 90(1): 17-24.
J. Halverson, L. Spelman. 2002. Sex determination and its role in management. in Komodo Dragons - Biology And Management, Smithsonian Institution Press, 2002. pp.165-177.
D. Eason, C. Millar, A. Cree, J. Halverson, D. Lambert. 2001. A comparison of five methods for assignment of sex in the Takahe (Aves:Porphyrio Mantelli). Journal of Zoology, 253(3):281-292.
C. Millar, G. Taylor, P. Moore, J. Halverson, D. Lambert. A novel restriction fragment length polymorphism for petrels or tube-nosed birds. Molecular Ecology 9(11):1915-1917.
A. Carmichael, A. Fridolfsson, J. Halverson, H. Ellengren. 2000. Male-biased mutation rates revealed from z and w chromosome-linked ATP synthase alpha-subunit (ATP5a1) sequences in birds. Journal of Molecular Evolution, 50(5): 443-447.

G. Shutler, P. Gagnon, G. Verret, H. Kalyn, M. Korkosh, E. Johnston, J. Halverson. 1999. Removal of a PCR inhibitor and resolution of DNA STR types in mixed human-canine stains from a five year old case. Journal of Forensic Science 44(3) :623-626. 1999.

J. Halverson, J. Dvorak, T. Stevenson. 1995. Microsatellite sequences for canine genotyping. United States Patent Application, Registration No. 36,377.
J. Halverson, G. Sverlow. 1994. Methods and compositions for the diagnosis of flea allergies. United States Patent Application, Serial No. 08/333,952.
J. Halverson, J. Dvorak. 1993. Avian sex identification probes. United States Patent Serial No. 08/194,131.
A. Fridolfsson, H. Cheng, N. Copeland, N. Jenkins, Liu Hsiao-Ching, T. Raudsepp, T. Woodage, C. Bhanu, J. Halverson, H. Ellengren. 1998. Evolution of the avian sex chromosomes from an ancestral pair of autosomes. Proceedings of the National Academy of Sciences of the United States Of America, 95(14):8147-8152.
C. Millar, C. Reed, J. Halverson, D. Lambert. 1997. Captive management and molecular sexing of endangered avian species: an application to the Black Stilt (Himantopus novaezelandiae) and hybrids. Biological Conservation 82(1):81-86.
T. Fleming, J. Halverson, J. Buchanan. 1996. Use of DNA analysis to identify sex of Northern Spotted Owls (Strix Occidentalis Caurina). Journal Of Raptor Research 30(3):118-122.
C. Millar, D. Lambert, S. Anderson, J. Halverson. 1996. Molecular sexing of the communally breeding Pukeko - an important ecological tool. Molecular Ecology5(2):289-293.
T. Sabo, R. Kesseli, J. Halverson, I. Nisbet, et al.1994. PCR-based method for sexing Roseate Terns (Sterna Dougallii). Auk111(4):1023-1027.
J. Halverson, J. Dvorak. 1993. Genetic control of sex determination in birds and the potential for its manipulation. Poultry Science72(5):890-896.
J. Dvorak, J. Halverson, P. Gulick, K. Rauen, et al. 1992. cDNA cloning of a Z-linked and W-linked gene in gallinaceous birds. Journal Of Heredity83(1):22-25.

Abstracts (Return to Top)
J. Halverson and C. Basten. The impact of canine inbreeding on likelihood calculations for the significance of DNA matches in forensic investigations. International Society of Animal Genetics. Tokyo, Japan. September 2004.
J. Halverson. 2002. Canine mitochondrial haplotyping and its use in criminal investigations. Cambridge Healthtech Institutes Fifth Annual DNA Forensics Meeting, Washington, Dc. June 2002.
J. Halverson, E. Johnston, L. Bickel, T. Sharp, J. Edwards. 2000. The AKC breed study: STR polymorphism at 17 loci in 91 AKC-registered breeds. International Society of Animal Genetics. Minneapolis, Minnesota, USA. July 2000.
J. Halverson, M. Malandro. 2000. The utility of mitochondrial hypervariable regions (Hv1 And Hv2) as forensic tools of feline and canine identification. Plant and Animal Genome Viii, San Diego, California. January 2000.
J. Halverson, J. Edwards. 1998. Microsatellite polymorphism in major dog breeds. International Society of Animal Genetics. Auckland, New Zealand. July 1998. Abstract in Animal Genetics 29 (Suppl.1)
J. Ziegle, Weller, J. Kuiper,M, M. Neff, J. Halverson, S. Bates. 1996. AFLP map of the dog genome. International Society of Animal Genetics. Tours, France. July 21-25, 1996. Abstract In Animal Genetics 27(Suppl.2)
D. Fantin, M. Bozzini, C. Gaiser, J. Halverson, S. Bates, J. Ziegle. 1996. Automating canine parentage and identification. International Society of Animal Genetics. Tours, France. July 21-25, 1996. Abstract in Animal Genetics 27(Suppl.2)

Presentations (Return to Top)
J. Halverson. 2005 Canine hair - linking victim to suspect. Forensic Hair Analysis Seminar, DePaul University, Chicago, IL. (invited)
J. Halverson, 2005. Hanging by a hair - animal trace evidence in forensic investigations. Annual Meeting of the American Academy of Forensic Science. New Orleans, LA.
J. Halverson, 2004. Hanging by a hair - animal-derived trace evidence in criminal investigation. Combined meeting of the Midatlantic Association of Forensic Scientists, the Midwestern Association of Forensic Scientists, the Southern Associations of Forensic Scientists, and the Canadian Society of Forensic Scientists. Orlando, FL.
J. Halverson, 2004. Animal DNA- a forensic tool. Animal forensics workshop at the biannual meeting of the International Society of Animal Genetics. Tokyo, Japan. (invited)
J. Halverson, 2004. Forensic DNA identification of feline hairs: Casework and a mitochondrial database. Annual Meeting of the American Academy of Forensic Science. Dallas, TX.
J. Halverson, A. Perroud, C. Gaiser. 2003. Forensic identification of canine hairs: is heteroplasmy significant? Promega 14th International Symposium on Human Identification. Phoenix, AZ October 2003 (Invited)
J. Halverson, N. Dvorak, A. Carrera, A. Kahn, A. Basten, C. Gaiser. Canine DNA testing and its use in criminal investigations. 2003. Bi-Annual Meeting of the European Academy of Forensic Science, Instanbul, Turkey September 2003. (Invited)
J. Halverson. 2002. Animal DNA and its application to homicide investigations. Virginia Homicide Investigators, September 30-October 2, 2002, Williamsburg, VA (Invited)
J. Halverson. 2000. DNA typing of animal-derived evidence in human crimes. Cambridge Healthtech Institute's Fourth Annual DNA Forensics , May 31-June 2, 2000. Springfield, VA (Invited).
J. Halverson. 1997. DNA testing in animals-from discovery to high-throughput application. Cambridge Healthtech Institute's Impact of Molecular Biology on Animal Health and Production Research, March 10-11, 1997. Baltimore, MD (Invited).

Continuing Education (Return to Top)
Forensic Hair Analysis Seminar, DePaul University, Chicago, IL. March, 2005.
Annual Meeting of the American Academy Of Forensic Science. New Orleans, LA. February 2005.
Combined meeting of the Midatlantic Association of Forensic Scientists, the Midwestern Association of Forensic Scientists, the Southern Associations of Forensic Scientists, and the Canadian Society of Forensic Scientists. Orlando, FL. September, 2004.
Hair Examination for DNA Analysts Workshop. Combined meeting of the Midatlantic Association of Forensic Scientists, the Midwestern Association of Forensic Scientists, the Southern Associations of Forensic Scientists, and the Canadian Society of Forensic Scientists. Orlando, FL. September, 2004.
International Society of Animal Genetics. Tokyo, Japan. September, 2004.
FBI DNA Auditor Workshop. Combined meeting of the Midatlantic Association of Forensic Scientists, the Midwestern Association of Forensic Scientists, the Southern Associations of Forensic Scientists, and the Canadian Society of Forensic Scientists. Orlando, FL. September, 2004.
Annual Meeting of the American Academy Of Forensic Science. Dallas, TX February 2004.
Promega 14th International Symposium on Human Identification. Phoenix, AZ October 2003.
Biannual Meeting of the European Academy of Forensic Science, Istanbul, Turky. September 2003.
Annual Meeting of the Association of Avian Veterinarians, Pittsburgh, PA. August 2003.
Annual Meeting of the American Academy Of Forensic Science. Chicago, ILl. February 2003.
Annual Meeting of the Association of Zoo Veterinarians, Milwaukee, WS. October 2002.
Annual Meeting of the Association of Avian Veterinarians, Monterey, CA. August, 2002.
DNA Forensics (5th Annual Meeting), Cambridge Healthtech Institute, Washington, DC. June 2002.
Biannual Meeting of the International Society of Animal Genetics. Minneapolis, MN, USA. July 2000.
DNA Forensics (4th Annual Meeting), Cambridge Healthtech Institute, Springfield, VA. May 2000.
Plant And Animal Genome Viii Conference, San Diego, CA. January 2000.
Plant And Animal Genome Vii Conference, San Diego, CA. January 1999.
Canine Health Foundation/American Kennel Club Canine Genetic Health Conference, St. Louis, MO. November 1999.
Biannual Meeting of the International Society of Animal Genetics. Auckland, New Zealand. July 1998.
Canine Health Foundation/American Kennel Club Molecular Genetics and Canine Genetic Health Conference, St. Louis, MO. November 1997.
Canine Health Foundation/American Kennel Club National Parent Club Canine Health Conference, St. Louis, MO. October 1997.
Impact Of Molecular Biology on Animal Health and Production Research, Cambridge Healthtech Institute, Baltimore, MD. March 1997.
Biannual Meeting of the International Society of Animal Genetics. Tours, France. July 1996.
American Kennel Club Parent Club Genetics Conference, Florham Park, NJ. October 1995.
American Kennel Club Parent Club Genetics Conference, Florham Park, NJ. October 1994.

Closed Casework (Return to Top)
State of Illinois v. Cecil Sutherland, 2004
Sutherland was retried for the abduction, rape and murder of 10-year old Amy Shultz. He had been convicted before but had appealed based on an incompetent defense. Mitochondrial typing of a human hair found on Amy was matched to Sutherland and excluded the other suspect in the case. Numerous dog hairs found on Amy's clothing matched Sutherland's dog Babe. The frequency of Babe's haplotype in QuestGen's database is 2.6% and the resulting likelihood ratio is 34.3.
Commonwealth of Pennsylvania v. Patrick Ramsey, 2003
An elderly woman, Mildred Hauser, was sexually assaulted and murdered in her home. Four dog hairs were recovered at the crime scene. Patrick Ramsay was arrested after using Hauser's credit card; some of her jewelry was in his possession. A STR profile (13 loci with allelic dropout in one locus) of one of the hairs matched a dog living in Ramsay's household. The likelihood ratio for the match was 1.4X 1010 and very strongly supported the prosecution's case that someone from Ramsay's household was involved in Hauser's murder. The District Attorney, Francis Schultz wrote "…we were able to show that the killer of Mildred Hauser was connected to the residence in Pittsburgh where the dog lived… The dog hair evidence was an important piece of evidence that boxed the defendant into a corner." Ramsay pled guilty to first degree murder and was sentenced to life in prison.
Regina v. Daniel McGowan, Leeds, United Kingdom, 2003.
Brian Keating was beaten and abducted from his home while his family was held at gun point. He was found beaten to death the next day. Numerous dog hairs were found on his clothing. The West Yorkshire Police believed that he had been transferred during the night to a van owned and driven by Daniel McGowan. A STR profile with 16 STR loci and minor allelic dropout matched the dog owned by Daniel McGowan. The likelihood ratio for the match was 5.3 X 1014 and very strongly supported the prosecution's case that McGowan had been involved in the abduction and murder. He and three other defendants were convicted.
State of Iowa v. Ben O'Donnell, 2003
Tracy Carson disappeared following a birthday party in a local bar. Her body was found 6 months later; she had been strangled, wrapped in a bolt of fabric, partially burned and then buried in a creek bed. Spring flooding unearthed her body. Examiners noted animal hairs on the fabric. Ben O'Donnell had "partied" with Tracy the night she disappeared and was the primary suspect. Police conjectured that the fabric wrapping the body had been taken from the home of O'Donnell's grandmother. Mitochondrial haplotypes of cat hairs found on the fabric (a common type with a frequency of 28.5% and an uncommon type with a frequency of 1.3%) did not exclude the three household cats. These frequencies are independent and can be combined to produce a likelihood ratio of 239. Other evidence was developed when investigators used luminol to find blood traces with matching DNA profiles to Tracy Carson in the truck of O'Donnell's car. He pleaded guilty to second-degree murder.
State of Florida v. Brent Robert Huck, 2003
April Misty Morse was kidnapped and murdered. Her body, bound with duct tape, was found in a coastal river. Several dog hairs were found on the duct tape. Police suspected an ex-boyfriend, Brent Robert Huck, had murdered her on his boat. The mitochondrial haplotype of the dog hairs matched that of Huck's dog. The frequency of the type was common at 10.4%; the likelihood ratio is 8.5. Brent Huck was convicted on both counts of kidnapping and murder.
Dog Attacks in Cook County, 2003
A woman was attacked and badly injured by two dogs in a park in Cook County, Illinois. Police searching for the dogs found a second female victim who died within a few hours. A dog approached the police threateningly was killed; tissue from the second victim was found in its stomach. Police trapped a number of dogs, including second dog whom they strongly suspected based on the description of the surviving victim. In order to reassure the public, police needed to confirm that this dog was the second attacker. Dr. Halverson tested the clothing of the two victims for canine DNA. Mitochondrial types from both dogs, along with the types of two dogs owned by the victims, accounted for all DNA profiles found on the clothing.
People vs. Jeanette Slover, Michael Slover Sr., and Michael Slover Jr.
A young woman, Karyn Hearn, disappeared from Decatur, Illinois. A few weeks later, local fisherman recovered trash bags from a local lake; the bags contained her dismembered body. The police found a dog hair on the tape used to close the bags. The police theory of the crime was that the family of her ex-husband, Michael Slover Jr., had murdered her because she planned to move to another state with their young son. The dog hair from the tape was tested by mitochondrial and STR typing and compared to a dog owned by the Slovers. The mitochondrial type matched and all alleles seen in a partial DNA profile from the hair were consistent with the defendants' dog. The defendants were convicted.
Shasta County, California v. Joshua Lee Davis, 2002.
The defendant was charged with animal cruelty for the vicious stabbing death of a dog. A STR profile (9 loci) from a knife in the defendant's possession did not exclude the dead dog. The likelihood ratio for the match was 2.5 X 108 and very strongly supported the prosecution's case that the suspect had killed the dog. The defendant pleaded guilty to one felony count of cruelty to animals.
Commonwealth of Pennsylvania v. Stephen Treiber, 2002.
The 2-year old daughter of Stephen Treiber died in a house fire in Mill Creek, Pennsylvania. During the investigation of the fire, Treiber showed police a threatening letter made by gluing words cut from newspapers to a sheet of paper. The trace evidence examiner noticed a dog hair encased in the glue with just the root end protruding. Alleles from a partial STR profile (7 loci with significant allele dropout) from the hair did not exclude one of Treiber' dogs. Ironically, this dog also died in the fire but its body was not consumed. The likelihood ratio for the match was 575 and strongly supported the prosecution's case that Treiber had fabricated the letter. He was convicted on all counts.
State of California v. David Westerfeld, 2002.
Seven year-old Danielle Van Dam was abducted from her home and murdered. Her body was found two weeks later. Danielle had been very close to the family's Weimeraner dog. The police suspected the Van Dam's neighbor, David Westerfeld, and searched his house and motor home. The mitochondrial type from dog hairs found in the motor home, on a quilt, and in the lint-trap of his dryer matched the Van Dam's dog. The frequency of the type was common at 9%; the likelihood ratio is 9.9. This was the first trial in the United States to admit canine mitochondrial DNA analysis as evidence. David Westerfeld was convicted on all counts.
People vs. Laykham, Ventura County, California 2002.
A small dog's determined barking alerted his elderly owner to the presence of an intruder. The suspect, Soum Laykham, attempted to rape the victim and fight off the Shih Tzu dog. A telephone call caused the suspect to flee. Dog hairs found on the suspect's clothing and in his closet matched the victim's dog by both mitochondrial typing and STR typing (9 loci). The likelihood ratio for the match was 7.7 X 108 and very strongly supported the prosecution's case that the hairs on the suspect's clothing had come from the victim's dog.
OCDA File #01F08374 People v. Richard David Ramirez, Javen Anthony Cervantes, and Stanley Cruz, Orange County, California, 2001.
The defendants were convicted of murdering Guillermo Carvajal, a resident of a rival gang area. His body was found wrapped in a blanket with numerous dog hairs. A STR profile of 12 STR loci with some allelic dropout matched a dog owned by one of the defendants. The likelihood ratio for the match was 3.8 X 107 and very strongly supported the prosecution's case that the defendant had been involved in the murder. Defense counsel stipulated to the canine DNA evidence and the defendants were sentenced to life in prison without the possibility of parole.
City of Tulsa vs. Michael Ohman & Vanessa Sorja, 2000.
Two pitbull-like dogs attacked an elderly woman, Wanda Cox, in her front yard after an early morning walk. Neighbors came to her aid and beat the dogs away. She was seriously injured in the attack and was permanently handicapped as a result. Despite the presence of several witnesses, the owners, Michael Ohman and Vanessa Sorja, claimed their dogs had not been the attackers. These same dogs had been involved in previous incidents but elderly witnesses had misidentified similar dogs from photos. STR profiles (10 loci) from hair from the victim's socks matched the defendants' dog. The likelihood ratio for the match was 1.4 X 109 using the Zoogen database (n = 558) and 2 X 108 using allele frequencies from a small pit bull database (n =24) and very strongly supported the prosecution's case that the hair came from the suspects' dog. They were found guilty of harboring a vicious animal.
New Jersey Division of Fish and Game Case#4-244-99/#5-244-99, 1999.
The tiger was alleged to have escaped from a private facility in New Jersey; the owner of the facility claimed the tiger had come from a nearby safari park. The evidence sample was a clump of fur found within the perimeter fence of the private facility. Domestic cat STR markers were used to profile the case samples and 20 tiger samples provided by Dr. Stephen O'Brien as a small database for the case. The clump of fur matched the deceased tiger. The facility has lately been closed and the tigers moved to a reserve in Texas.
Twelfth Judicial District State of New Mexico. State vs. Charles Martinez (Cause No. CR-99-108) and State vs. Chris Faviel (Cause No. CR-98-64), 1998-1999.
A woman's body was found in the New Mexico desert. The woman had been harassing a former boy friend and police suspected that the ex-boyfriend and his new lover had murdered her. A partial canine STR profile (8 loci with some allele dropout) from a single hair found on a sock of the victim did not exclude the dog owned by one of the suspects. The likelihood ratio for the match was 7.6 X 103 and very strongly supported the prosecution's case that the hair came from the suspect's dog. The lover, Charles Martinez, was tried and convicted. The ex-boyfriend, Chris Faviel, pled guilty.
Office of the Prosecuting Attorney of King County Washington, 1998. The State Vs, Leulualahi (97-C-08256-9) and State vs. Tuilefano (97-C-01391-3) for the murder of Jay Johnson and Raquel Rivera.
Gang members looking for drug money murdered a young couple during a home invasion. The couple's dog Chief attacked the invaders at the door and was shot twice at short range; he died later in surgery. The suspects were still wearing their blood-spattered clothing when arrested later that day. Canine STR profiles (10 loci) from bloodstains found on the clothing of the suspects did not exclude Chief. The likelihood ratio supporting the prosecution's hypothesis that the blood spatters were from Chief was 4.8 X 109. The use of the canine DNA results was successfully appealed in 2003 based on the lack of an admissibility hearing and the lack of peer-reviewed publications on the canine STR markers used at that time of the appeal. Since then, several reports have been published in peer-reviewed forensic journals.
Tom O'Connor vs. Irish Coursing Club, 1998.
Dr. Halverson worked with Pat Dalton, a prominent breeder of racing greyhounds, to vindicate Tom O'Connor of misrepresentation in the registration of a litter of puppies. In 1993, Serology Ltd., the Club's DNA testing laboratory, had purported that one puppy in the litter was excluded to the dam. DNA testing by four other laboratories worldwide, including Zoogen, Inc. had found no evidence of exclusion. Dr. Halverson testified at the 1998 hearing. Serology LTD was found to be in error. Tom O'Connor was awarded substantial financial compensation.
Royal Canadian Mounted Police Detachment File Number 91-0693 (reference: Schraeder, Daniel Fred-Murder of), 1996-1997.
Daniel Schraeder, an elderly man in Vernon, British Colombia, was murdered by blunt force trauma along with his small dog. Mixed-specie bloodstains on the suspect's jeans were not amenable to the RFLP methods available at the time. The case was reopened in 1996 and canine STR profiles from the non-human blood on the jeans matched the victim's dog. The evidence was used by the Canadian Royal Counsel to assign murder charges against the primary suspect. In September 1997, the suspect filed a guilty plea.

Contact Information

Joy Halverson, DVM, MPVM
QuestGen Forensics
1046 Olive Drive Suite1
Davis, CA 95616
Phone: (530) 758-4254
Fax: (530) 750-5758
Email address: halverjl@hotmail.com

 

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